El arquetipo vertebrado delineado por Richard Owen a mediados del Siglo XIX fue un modelo concebido para pautar el análisis morfológico de los seres vivos dentro de los lineamientos definidos por la idea de una unidad de composición cuando ella se aplicaba al caso específico de los vertebrados. Pero aquí, además de analizar la razón de ser originaria de ese arquetipo y su modo de operar dentro del contexto epistémico en el que efectivamente surgió, también aludiremos al modo en que (...) ese modelo pudo pasar a ser considerado, no ya como una idealización, o una abstracción metodológicamente operativa, sino como una aproximación a una entidad real que debía ser individualizada y debidamente descripta. (shrink)

Un vínculo necesario entre estética y socialización explica la profundidad sensible que debe construir todo cambio de paradigma y toda re-socialización. La democracia no funciona sin una apropiación individual y colectiva del sentir y de la producción de sentir. Es a partir entonces del trabajo artístico y de la instrospección y exteriorización de la dimensión sensible, que toda transformación puede ser aplicada. Se recorren parcialmente los pensamientos de Alain Badiou y Bernard Stiegler. A required link between aesthetic and socialization explains (...) the sensitive depth that all paradigm and re-socialization must build. Democracy does not work without an individual and collective ownership of feeling and feeling's production. It is from the artwork and the introspection and externalization of the sensitive dimension, that any transformation can be applied. The article covers partially the thoughts of Alain Badiou and Bernard Stiegler. (shrink)

Admitiendo la distinción propuesta por Mayr entre biología funcional y biología evolutiva, sugerimos que estos dos dominios de investigación siguen dos distintos modos de considerar lo viviente que pueden ser entendidos, pero en dos sentidos diferentes, como teleológicos. Para distinguir esas dos formas de teleología hacemos una comparación entre la explicación funcional típica de la biología funcional y la explicación seleccional propia de la biología evolutiva. Cada uno de estos tipos de explicación obedece a una regla metodológica especial: la explicación (...) funcional sigue al principio de adecuación autopoiética; y la explicación seleccional sigue al principio de adecuación adaptativa. Pero, mientras el primero será presentado como estando subordinado a un principio general da causación; el segundo será presentado como siendo independiente de él. Finalmente, en el contexto de una breve discusión relativa al concepto de symmorphosis, sostenemos que, en la biología contemporánea, la noción de adecuación adaptiva es preeminente sobre la noción de adecuación funcional: esta encuentra su fundamento en aquella.Admitting Mayr’s distinction between functional and evolutionary biology we suggest that these two dominions of inquiry follow two different modes of onsider the living that can be considered, but in two different senses, as being teleological. To distinguish these two forms of teleology we make a comparison between the functional explanation, typical of functional biology, and the selective explanation, proper of evolutionary biology. Each one of this kind of explanation obeys an special methodological rule: the functional explanation follows the principle of autopoietic adequacy; and the selective explanation follows the principle of adaptive adequacy. But, while the former principle will be presented as being subordinated to a general principle of causation; the later will be presented as being independent of it. Finally, in the context of a brief discussion concerning the concept of symmorphosis, we argue that, in contemporary biology, the notion of adaptive adequacy is preeminent over the notion of functional adequacy: the later find its fundament in the former. (shrink)

En sus Principios de biología de 1864, Spencer esboza una complementación entre el cuvierianismo transformacional mitigado que daba sentido a la idea de equilibración directa ahí presentada, y la teoría de la selección natural que Darwin ya había formulado en 1859. Era a este último mecanismo que Spencer denominaba "equilibración indirecta". Según Spencer, esta segunda forma de equilibración permitía explicar fenómenos evolutivos que la primera, la equilibración directa, no podía causar; aunque para él también era evidente que el accionar de (...) esa segunda equilibración debía siempre subordinarse al control de la primera. In his Principles of Biology, published in 1864, Spencer outlines a complementarity between the mitigated transformational cuvierianism that was in the basis of his idea of direct equilibration, and the theory of the natural selection that Darwin had already formulated in 1859. Spencer called the mechanism of natural selection indirect equilibration. According to him, this second form of equilibration enabled the explanation of some evolutionary phenomena that could not be caused by direct equilibration. However, for Spencer, it was clear that the operation of this second equilibration had to be always subordinated to the control of the first one. (shrink)

Las presiones selectivas no son factores ambientales que existan independientemente de los linajes por ellas afectados. Su existencia y su configuración siempre dependen de las alternativas de evolución que el propio linaje genera y acepta; y dicha configuración, necesariamente, se va alterando conforme el propio linaje evoluciona. Aclarar eso es muy importante para desestimar algunas reticencias generadas por la dicotomía próximo-remoto. Esta polaridad continúa siendo un recurso conceptual válido e insustituible para entender la especificidad de la Biología Evolucionaria. Enmendar algunos (...) equívocos generados por Mayr en su caracterización del concepto de causa remota es una alternativa mejor que la de pensar en abandonar o en relativizar esa distinción. (shrink)

La distinción propuesta por Ernst Mayr y François Jacob entre biología funcionaly biología evolutiva es una clave fundamental para un tratamiento claro y concluyente de dos problemas que, recurrentemente, se colocan en el ámbito de la Filosofía de la Biología: aquel que atañe al posible carácter teleológico de las explicaciones biológicas y aquel relativo a la autonomía de la Biología en relación a la Física y a la Química. Sin sostener una alternativa de solución para ninguno de estos problemas, presentamos (...) esa oposición mostrando su relevancia para el tratamiento de tales cuestiones. Nuestra idea es que las mismas no deberían plantearse sin especificar a cual de los dos dominios de la ciencias de la vida nos estamos refiriendo.Mayr’s and Jacob’s distinction between functional biology and evolutionary biology is a fundamental key for a clearer and more conclusive treatment of two problems that, currently, are discussed in Philosophy of Biology: that which matter to the teleological character of biological explanations and that which matter to the autonomy of Biology from Physic and Chemistry. Without hold any alternative solution for none of these problems, we present that opposition showing its relevance for the treatment of both questions. Our idea is that these problems shouldn’t be posed without a specification of the domain of Biology that is being considered. (shrink)

RESUMEN Se analiza si la versión de la justicia como equidad, presentada en El liberalismo político, es genuinamente una concepción política. Se examina el problema de la razonabilidad de las doctrinas comprehensivas, y se indaga luego si el argumento en dos etapas afecta la integridad estructural del liberalismo político. Se concluye que J. Rawls fracasa en su intento de justificar un liberalismo independiente de una doctrina comprehensiva de carácter liberal. ABSTRACT The article analyzes whether the conception of justice as fairness, (...) set forth by Rawls in Political Liberalism, is a genuine political conception. It first examines the issue of reasonableness of comprehensive doctrines and then goes on to inquire whether the two-step argument affects the structural integrity of political liberalism. The paper concludes that J. Rawls fails in his attempt to justify liberalism independently of a comprehensive liberal doctrine. (shrink)

Asumiendo y profundizando la distinción entre Biología Funcional y Biología Evolutiva conforme ha sido propuesta por Ernst Mayr y François Jacob, comparamos la naturaleza de los procedimientos experimentales que se realizan en uno y otro dominio de las ciencias de la vida. En vistas a esto, introducimos ciertas precisiones sobre las nociones de observación y experimento e intentamos elucidar los presupuestos sobre los que se apoya la experimentación en biología funcional. A continuación, presentamos los fundamentos y las particularidades de los (...) procedimientos experimentales desarrollados en el contexto de ese capítulo de la biología evolutiva que es la genética de poblaciones, y analizamos las peculiaridades de las variables sobre la que estos se ejercen. Esto nos permite mostrar la diferencia radical que existe entre operar sobre variables poblacionales y operar sobre variables fisiológicas.In this article we compare the nature of the experimental procedures that are carried out in different areas of biology. This is done by accepting and deepening the distinction between Functional and Evolutive Biology as it is argued for by Ernst Mayr and François Jacob. Keeping this in mind, we introduce some remarks on the notions of observation and experiment and try to clear up the presuppositions on the basis of which experimentation in Functional Biology rest. In the wake of it, we show the foundations and the main aspects of those experimental procedures that are developed in the context of that chapter of Evolutive Biology called Genetics of Populations and analyze the peculiarities of the variables by which those experiments are carried out. This allows us to show the radical difference between working on population and physiological variables. (shrink)

Contrariando la concepción etiológica de las funciones biológicas, se pretende mostrar que las explicaciones por selección natural, lejos de justificar atribuciones funcionales, las presuponen; y que estas atribuciones pueden ser entendidas apelando a una especificación particular de la concepción s..

Cuvier’s confidence in the possibility of reconstructing the way of life of an extinct animal from a complete knowledge of its anatomy and its physiology could make us think that his program for paleontology somewhat anticipated Darwinian adaptacionism. Nevertheless, even if this interpretation seems reasonable, it ignores the cognitive aims and the fundamental presuppositions of the Cuvierian program. This ignorance conspires against a correct evaluation of the meaning and the impact of the results obtained by Cuvier in the study of (...) fossils. The relevance of those results is in showing that extinct living beings were subordinated to the same laws of organization as living beings existing today. (shrink)

Taking as a starting point Brandon's account of the principle of natural selection, we argue that it is possible to consider such a principle as bearing the same status of the principle of causation, to wit, that of a methodological rule whose function would be to introduce a "teleological mode of inquiring the living". This way of understanding the principle of natural selection will drive us into an interpretation of Darwinism that is close to that one argued for by Daniel (...) Dennett. (shrink)

In opposition to the etiological conception of biological functions, this paper attempts to show that explanations by natural selection, far from justifying functional attributions, presuppose them, and that these attributions may be understood by appealing to a particular specification of the systemic conception of function, that is, the biological conception of function. This paper argues that the etiological conception of function is based on two fundamental errors: confusing the concept of function with the concept of adaptation, and confusing selectional explanations (...) with functional analyses. This explains the inappropriate demand that those analyses include an etiological explanatory content and make possible the distinction between adaptations and exaptations. (shrink)

En la Teoría de la Selección Natural, el concepto de función biológica debe suponerse para delimitar el concepto de aptitud; y éste debe suponerse para delimitar el concepto de adaptación y también para explicar el fenómeno al que este último alude. Esos tres conceptos, por otra parte, son especificaciones de tres conceptos de aplicación más universal. El concepto de función biológica es un caso particular del concepto general de función; y el concepto de aptitud especifica el concepto de eficiencia. El (...) concepto de adaptación, mientras tanto, es una especificación del concepto de diseño. La visualización de esas articulaciones conceptuales permite entender la legitimidad de la naturalización de la teleología operada por el darwinismo.In Natural Selection Theory, the concept of biological function must be supposed to elucidate the concept of fitness; and this one must be supposed to elucidate the concept of adaptation and also to explain the phenomenon to which the last one alludes. These three concepts, on the other hand, are specificationsof three concepts of more universal application. The concept of biological function is a particular case of the general concept of function; and the concept of fitness is a specification of the concept of efficiency. The concept of adaptation, meanwhile, specifies the concept of design. The visualization of these conceptualjoints allows to understanding the legitimacy of the naturalization of the teleology produced by the Darwinism. (shrink)

El gran malentendido que está por detrás de la concepción etiológicadel concepto de función es haber confundido este último con el concepto de adaptación. Las explicaciones por selección natural no justifican imputaciones funcionales: ellas explican la configuración de determinada estructura orgánic..

The Darwinian revolution supposed and imposed a much broader and more complex concept of environment than that which, until that moment, had been considered by most as part of natural history. Until Darwin, the environment of living beings had been regarded, almost exclusively, as the inanimate surroundings. This pre-Darwinian notion of environment included physicochemical and climatic variables: the living beings themselves were scarcely considered, or they were regarded just as food to be assimilated. In contrast, with Darwin, the influence of (...) living surroundings began to be regarded as having decisive importance for understanding the distribution, configuration, and viability of living beings. The relationships established among different living beings became the main and more significant features of the biological environment. This conceptual change was an exigency derived from the formulation and justification of the theory of natural selection. The idea of struggle for existence supposed by that theory requires that the biotic environment be considered critical in determining the viability of each living being. (shrink)

En la Teoría de la Selección Natural, el concepto de función biológica debe suponerse para delimitar el concepto de aptitud; y éste debe suponerse para delimitar el concepto de adaptación y también para explicar el fenómeno al que este último alude. Esos tres conceptos, por otra parte, son especificaciones de tres conceptos de aplicación más universal. El concepto de función biológica es un caso particular del concepto general de función; y el concepto de aptitud especifica el concepto de eficiencia. El (...) concepto de adaptación, mientras tanto, es una especificación del concepto de diseño. La visualización de esas articulaciones conceptuales permite entender la legitimidad de la naturalización de la teleología operada por el darwinismo.In Natural Selection Theory, the concept of biological function must be supposed to elucidate the concept of fitness; and this one must be supposed to elucidate the concept of adaptation and also to explain the phenomenon to which the last one alludes. These three concepts, on the other hand, are specificationsof three concepts of more universal application. The concept of biological function is a particular case of the general concept of function; and the concept of fitness is a specification of the concept of efficiency. The concept of adaptation, meanwhile, specifies the concept of design. The visualization of these conceptualjoints allows to understanding the legitimacy of the naturalization of the teleology produced by the Darwinism. (shrink)

Although the knowledge about biological systems has advanced exponentially in recent decades, it is surprising to realize that the very definition of Life keeps presenting theoretical challenges. Even if several lines of reasoning seek to identify the essence of life phenomenon, most of these thoughts contain fundamental problem in their basic conceptual structure. Most concepts fail to identify either necessary or sufficient features to define life. Here, we analyzed the main conceptual frameworks regarding theoretical aspects that have been supporting the (...) most accepted concepts of life, such as (i) the physical, (ii) the cellular and (iii) the molecular approaches. Based on an ontological analysis, we propose that Life should not be positioned under the ontological category of Matter. Yet, life should be better understood under the top-level ontology of “Process”. Exercising an epistemological approach, we propose that the essential characteristic that pervades each and every living being is the presence of organic codes. Therefore, we explore theories in biosemiotics and code biology in order to propose a clear concept of life as a macrocode composed by multiple inter-related coding layers. This way, as life is a sort of metaphysical process of encoding, the living beings became the molecular materialization of that process. From the proposed concept, we show that the evolutionary process is a fundamental characteristic for life’s maintenance but it is not necessary to define life, as many organisms are clearly alive but they do not participate in the evolutionary process (such as infertile hybrids). The current proposition opens a fertile field of debate in astrobiology, epistemology, biosemiotics, code biology and robotics. (shrink)

Según Ron Amundson, los actuales desarrollos de la biología evolucionaria del desarrollo podrían estar exigiéndonos una revisión de la distinción entre causas próximas y remotas de los fenómenos biológicos propuesta por Ernst Mayr. En este trabajo argumentaré que, aunque sea cierto que la dicotomía ..

Research in modern biology has largely been developed according to two main ways of inquiry, as they were outlined by Charles Darwin and Claude Bernard. Each stands for a specific approach to the living corresponding to two different methodological rules: the principle of natural selection and the principle of causation.

La idea de herencia extendida se ha transformado en un leitmotiv de lo que se ha dado en llamar ‘síntesis extendida’. Dicha idea, entretanto, lleva a ignorar una de las marcas distintivas de lo que clásicamente se ha entendido por transmisión hereditaria: su estrecha asociación con el proceso de reproducción. Sin considerar tal asociación, la noción de herencia, más que ampliarse, de hecho se disuelve; y aceptar esa disolución acarrearía demasiadas dificultades para el discurso de las ciencias biológicas. Una de (...) esas dificultades tiene que ver con el concepto de variación hereditaria. Con la disolución de la noción de herencia, ese concepto también se disolvería; y, sin éste, la Teoría de la Selección Natural no puede articularse. Pero, no habiendo motivos valederos para aceptar semejante costo teórico, aquí quiero mostrar que ambos conceptos, el de herencia y el de variación hereditaria, pueden y deben ser elucidados sin aceptar la idea de herencia ampliada. (shrink)

Having as starting point that a proximal cause is one which effects can be registered in the states of an individual organism, in this work I will argue that what defines an ultimate cause is the fact that its effects can be registered in the evolution of lineages, and not simply in population states. This, on the other hand, not only will allow us to clarify how the developmental constraints can be understood as causes of the evolutionary phenomena; but also (...) it will allow us to point the eminently historical character of Developmental Evolutionary Biology that Wallace Arthur lessened in Biased Embryos and Evolution. Thus, and following this latter reasoning, I will also try to show that, although the action of developmental constraints could happen beyond the limits of a population or a particular species, that does not imply that we are before a comeback of typological thought in Evolutionary Biology. (shrink)

The main cognitive target of Ecology is the functional analysis of the ecological processes and systems. It does not suppose, meanwhile, that these processes and systems are designed systems and processes like individual leaving beings. The Ecology, likewise Physiology, is constitutively guided by the presupposition of a privileged state , to be explained, that it is the persistence of the systems and processes that she studied; and its functional analyses obey to this presupposition. Ecology supposes an ideal of natural order (...) , under which consideration this privileged state is thinking as an improbable state of things and, for that, needed of explanation; and, in what concerns this latter point, the analogy between Ecology and Physiology can also result instructive. DOI:10.5007/1808-1711.2010v14n3p349. (shrink)

RESUMEN: La idea de selección interna, propuesta originalmente por Lancelot Whyte, no sólo sirve para entender el papel causal que los constreñimientos del desarrollo tienen en evolución; sino que además puede hacernos comprender de qué modo esos factores organísmicos o internos, cuya importancia la Biología Evolucionaria del Desarrollo hoy quiere rescatar, son pasibles de ser considerados desde una perspectiva variacional o seleccional compatible, pero no asimilable, a la Teoría de la Selección Natural. Así, considerado como un concepto autónomo y diferente (...) al de selección natural, el concepto de selección interna puede permitirnos comprender de qué modo eso que Wallace Arthur llama developmental bias constituye una causa, efectiva y positiva, de transformacionesevolutivas irreversibles y acumulables.ABSTRACT: The idea of internal selection, originally proposed by Lancelot Whyte, not only helps to understand the causal role that the developmental constraints have in evoltion, but it can also show how these organismic or internal factors, which relevancy the Evolutionary Developmental Biology remarks today, can be considered from a variational or selectional perspective that is compatible with, but not reducible to, Natural Selection Theory. Thus, being considered as a concept autonomous and different from natural selection, the concept of internal selection can allow us to understand how that phenomenon that Wallace Arthur calls developmental bias constitutes an effective and positive cause of irreversible and cumulative evolutionary changes. (shrink)

En los últimos años, algunos autores han venido sosteniendo que la validez de la representación arborescente del patrón filético generado por la evolución está siendo menoscabada por el reconocimiento del impacto evolutivo que tendrían la transferencia genética horizontal, la simbiogénesis, la especiación por hibridación y la introgresión. Esa concepción o representación cladogenética de la evolución sólo nos dejaría ver un aspecto parcial de las relaciones de filiación que conectan a los diferentes linajes de seres vivos; ocultando otro aspecto cuya representación (...) más adecuada sería un reticulado de relaciones entre taxones. Con todo, aún sin poner en tela de juicio la frecuencia y la relevancia explicativa de dichos fenómenos, en este trabajo se intentará mostrar que su admisión y reconocimiento no tiene por qué ir en detrimento de esa concepción cladogenética de la evolución cuya relativización se está pregonando. Por el contrario, la comprensión y la representación de tales fenómenos supone dicha concepción. Palabras clave: Filogénesis, Especiación por hibridación, Introgresión, Simbiogénesis, TGH. (shrink)

Firstly, in this article it is examined the nature of the putative remarks concerning evolution of species that are found in the works that Richard Owen published before 1858; and then it is made the same thing with the few and vague evolutionist conjectures that Owen certainly made after the public presentation of Natural Selection Theory. Regarding the former topic, the goal will be to highlight the ambiguity of those Owen’s remarks, and concerning the latter topic what is looked for (...) is to show that, when he explained his transmutationist thesis, Owen didn’t go beyond a pious theistic evolutionism, without postulating any mechanism of evolutionary change and remaining faithful to Design Theology praised by Paley. (shrink)

The distinction between evolutionary and functional biology is a fundamental clue to a conclusive approach of one of the most controversial issues in the philosophy of biology: the possibility of reducing biology to physics and chemistry. The questions that defines the cognitive targets of functio..

Segundo Daniel Dennett insistiu em diferentes trabalhos, o programa adaptacionista darwiniano constitui uma legítima e insubstituível translação da perspectiva intencional ao domínio da biologia. Mas, para que essa tese possa ser formulada com toda clareza, e não fique no plano da simples metáfora – coisa que não é o objetivo de Dennett – é necessário esclarecer qual seria o sistema intencional cujo comportamento estudamos conforme essa perspectiva. Assim, e contra a alternativa escolhida pelo próprio Dennett, e retomando uma proposta do (...) Karl Popper, sustentaremos que esse sistema não é a natureza como um tudo, mas sim o sistema constituído por uma população ou uma linhagem de organismos: eles são os agentes dos processos adaptativos. Não é nossa meta, entretanto, contradizer as teses centrais da Dennett; mas sim as reforçar.How do species think? Daniel Dennett has insisted in different occasions that Darwin’s Adaptationist Program constitutes a legitimate and hardly displaceable translation of the intentional point of view in the Biology field. However, in order for us to be able to state this thesis clearly and to avoid getting into the domain of mere metaphor – which is not Dennett’s purpose -, we have to single out what intentional system the behaviour of which we study on such a perspective. That being noted, we shall argue, contrary to the alternative that Dennett himself picked out, and close to Karl Popper’s suggestion, that such a system is not nature as a whole, but the system made up by a population or a class of organisms: those are the agents of the adaptative processes. We do not, however, aim at contradicting Dennett’s overall strategy; rather, we aim at reinforcing it. (shrink)

The term ideology is usually used in three different senses, each of which alludes to a different concept: ideology in the sense originally proposed by Marx and Engels; ideology as a political thought, which ended up being the most frequently used notion; and ideology as a bias that affects the positions that can be adopted in the field of theory. This last notion will be analyzed here; trying to show its difference, but also its relationships, with the other two notions. (...) An attempt will also be made to establish a difference between the notions of ideological bias and of political position in theory. (shrink)